Forestry Commission publications
Bryce, J., Cartmel, S. and Quine, C. P. (2005). Habitat use by red and grey squirrels: results of two recent studies and implications for management (PDF-546K) . Forestry Commission Information Note 76. Forestry Commission, Edinburgh.
Mayle, B. A. (2004) Grey squirrel management in woodlands, in Managing woodlands and their mammals (Quine, C., Shore, R. & Trout, R., Eds.): proceedings of a symposium organised jointly by the Mammal Society and the Forestry Commission, 45-49, Forestry Commission, Edinburgh.
Gurnell, J., Lurz, P.W.W., Pepper, H. (2001) Practical techniques for surveying and monitoring squirrels (PDF-3830K) , Forestry Commission Practice Note 11, Forestry Commission, Edinburgh.
Pepper, H., Bryce, J., Cartmel, S. (2001), Squirrel management: 2. Co-existence of red squirrels and grey squirrels, in Forest Research Annual Report and Accounts 1999-2000. Forestry Commission, Edinburgh.
Pepper, H., Moore, H. (2001) Squirrel management: 1. Development of a contraceptive vaccine for the grey squirrel, in Forest Research Annual Report and Accounts 1999-2000. Forestry Commission, Edinburgh.
Pepper, H., Patterson, G. (2001) Red squirrel conservation (PDF-250K) , Forestry Commission Practice Note 5 (revised). Forestry Commission, Edinburgh.
Pepper, H. (1998) Nearest neighbour method for quantifying wildlife damage to trees in woodland (PDF-139K) , Forestry Commission Practice Note 1. Forestry Commission, Edinburgh.
Nichols, C.P., Gregory, N.G., Goode, N., Gill, R.M.A., and Drewe, J.A.(2017) Can the grey squirrel, Sciurus carolinensis, utilise calcium oxalate? – Implications for bark stripping behaviour Journal of Animal Physiology and Animal Nutrition. DOI: 10.1111/jpn.12740
Nichols, C.P. and Gill, R.M.A. (2016) Bark stripping behaviour by the grey squirrel,Sciurus carolinensis. Chapter 19 in: Shuttleworth, C. and Gurnell, J. (Eds) The Grey Squirrel. The European Squirrel Initiative.532p
Nichols, C.P., Gill, R.M.A., Drewe, J.A. Goode, N. Gregory, N. (2016) A novel causal mechanism for grey squirrel bark stripping: The Calcium hypothesis. Forest Ecology and Management. 367 12-20.
Mayle, B., Ferryman, M., Peace, A., Yoder, C., Miller, L., Cowan, D., (2013) The use of DiazaCon™ to limit fertility by reducing serum cholesterol in female grey squirrels, Sciurus carolinensis. Pest Management Science 69, 414-424.
Mayle, B.A. and Broome, A. (2013) Changes in the impact and control of an invasive alien, the grey squirrel Sciurus carolinensis, as determined from regional surveys. Pest Management Science, 69, 323-333.
Willoughby, I., Jinks, R.L., Morgan, G.M., Pepper, H., Budd, J. and Mayle, B.M. (2011). The use of repellents to reduce predation of tree seed by wood mice (Apodemus sylvaticus L.) and grey squirrels ( Sciurus carolinensis Gmelin) . European Journal of Forest Research. 130, 4, 601-611.
Yoder, C. A., Mayle, B.A., Furculow, C.A., Cowan, D,P. & Fagerstone, K.A. (2011). Feeding of grey squirrels (Sciurus carolinensis) with the contraceptive agent DiazaCon: effect on cholesterol, hematology, and blood chemistry. Integrative Zoology 6: 409-419.
Mayle, B. A., Proudfoot, J., Poole, J. (2009). Influence of tree size and dominance on incidence of bark stripping by grey squirrels to oak and impact on tree growth. Forestry 2009,82,4, 431-444. Full text , PDF (201K)
Ferryman, M., Mayle, B. A. and Morgan, G. W. (2006). Visual method for evaluating the state of sexual development in male grey squirrels ( Sciurus carolinensis). Reproduction, Fertility and Development 18, 383-393.
Mayle, B. A. (2005). Britain’s woodlands under threat; Grey squirrels and the risk they pose to European woodlands. Trees, Journal of the International Tree Foundation. 65 9-11.
Pepper, H.W., G. Kerr (2005). Preliminary studies on collars to protect trees from grey squirrel bark-stripping damage. Quarterly Journal of Forestry 99(2): 105-112.
Gurnell, J., Lurz, P.W.W., Shirley, M.D.F., Cartmel, S., Garson, P.J., Magris, L., Steele, J. (2004) Monitoring red squirrels (Sciurus vulgaris) and grey squirrels (Sciurus carolinensis) in Britain (PDF-347K) , Mammal Review. 34, 51-74.
Hewson, C., Fuller, R., Mayle, B., Smith, K. (2004) Possible impacts of grey squirrels on birds and other wildlife. British Wildlife 15, 183-191.
Mayle, B., Gurnell, J. (2004) Squirrel control in conifers, Forestry and British Timber, April 2004, 16-19.
Gurnell, J., Mayle, B.A. (2003) Ecological impacts of the alien grey squirrel (Sciurus carolinensis) in Britain, conference proceedings: MammAliens - a one day conference on the problems caused by non-native British mammals, 40-45, Mammals Trust UK.