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Ground Beetles (Carabidae) can be used to demonstrate changes in prey invertebrates. At Alice Holt the majority of beetles found are Abax parallelepipedus and Pterostichus madidus. Although variable year on year their populations have remained relatively stable. Species associated with dead wood and damp conditions (Cychrus caraboides, Nebrai brevicollis) have increased significantly most likely due to increases in litter accumulation (evident from the soil survey) dead wood, and shade associated with natural stand ageing.
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